Cambridgeshire - March 1939. As the clouds of war begin to gather in Europe, the Talbot family of rural Blackberry Farm will be torn apart, just as so many families all over the world will be. Life will never be the same again. Whilst in London, the Salmons family will feel the pain of parting and loss. Brought together by war, the two families become intertwined and, as the outlook looks bleak, they must draw on each other’s strength to fight through the hard times.

As war looms in Europe, 18-year-old Jane Shaw runs the household as an unpaid servant on a meagre budget. When tragedy strikes and she loses her beloved younger brother in suspicious circumstances, Jane takes a position in service looking after Ned, a troubled young boy. Here she meets dashing David Heron, Ned's Uncle and they form an immediate attachment until she is dismissed after being accused of stealing a piece of jewellery. Returning home, Jane and her sister, Melia, are soon orphaned and left at the mercy of their Aunt Alice.

Lady Joanna Harcourt and her daughter, Sarah, discover their life of luxury at Goodwill House is over. Because with Lord Harcourt away fighting, the Harcourt girls are facing financial ruin. Lady Joanna is terrified of losing their home, but for Sarah, this means her dreams of becoming a doctor are over. Headstrong Sarah isn’t one to quit! War or not, she’s determined to find a way to save her home and follow her dreams - whatever it takes.

When sixteen-year-old Sally is evacuated to the English south coast, she is terrified by what lies ahead of her. All she knows are the sights and sounds of London's East End – but Sally swallows her tears as they leave the familiar landmarks behind, knowing that she has to be a Grown-Up Girl and play mother to her six-year-old brother Ernie. Playing mother is nothing new for Sally – their real mother Florrie, a good-time girl, hasn't even come to the station to wave them off and Ernie, crippled at an early age by polio, is used to depending on his older sister.

The world is on the brink of war when Elizabeth Mowbray breaks her engagement with a tea planter in India and returns home to the English countryside. Desperate to escape a stifling life under her parents’ roof, she moves to London seeking adventure and excitement.

Following a humiliating experience involving the man she thought she’d marry, Iris Tredwick signs up to the Wrens in order to escape and - hopefully - find ‘the right sort’ of man to please her mother. After a bumpy start, Iris manages to befriend outspoken Mary and dreamer Sally as they are sent to their first posting - in Orkney. There she meets engineer Rob, whose flirtatious nature both charms and confounds straight-laced Iris. Much more appropriate for her is local doctor Stewart, if only she felt the same spark for him as she does for Rob.

Liverpool, 1939. Bella Rogers thought her life was just starting; she can’t wait to get married to her childhood sweetheart, Bobby. But when war is declared and Bobby is called up, his letters stop, leaving Bella heartbroken and alone. Then her family is torn apart by tragedy when her angelic 5-year-old sister dies from a terrible illness. They can barely afford a funeral, but the church is packed with the entire community, devastated for their loss. Grieving for her sister and lost love, Bella finds a job at the Bryant and May factory, making matches.

Maggie Bailey has not had an easy childhood. Her father, Michael, always too easy with his fists after spending the family’s rent money down the pub. Just sixteen, in love and blinded by promises, Maggie sees marriage to handsome Jack as her great escape. However, she soon finds herself abandoned with a beautiful baby when Jack disappears. Maggie is forced to seek a new life away from the East End of London and finds herself a job at a hotel in Yarmouth.

It’s 1942, and London remains under constant threat of enemy attack as the second world war rages on. In the Bethnal Green neighborhood, Nellie Morris counts every day lucky that she emerges from the underground shelters unharmed, her loving family still surrounding her. Three years into the war, she’s grateful to hold onto remnants of normalcy—her job as assisting the mayor and nights spent at the local pub with her best friend. But after a chance encounter with Ray, an American airman stationed nearby, Nellie becomes enchanted with the idea of a broader world.

Bristol, 1939. School leaver Maisie Miles suspects her father, a small-time crook, has an ulterior motive for insisting she gets a job at the W. D. & H. O. Wills tobacco factory but keeps it to herself. She's befriended by effervescent Phyllis Mason and kind-hearted Bridget Milligan who take pity on her and take Maisie under their wing. But beneath their happy-go-lucky exteriors, they all harbour dreams and worries about what the future holds.

Occupied Poland, 1942: Despite the constant presence of German soldiers in her village, Emilia is allowed certain freedoms as a midwife—the most precious is innocently cycling past Nazi checkpoints to the homes of expectant mothers on her rounds. But Emilia has a secret: for years she’s also been visiting the hidden Jewish mothers and working for the resistance…until she is betrayed.

1916. At the onset of war, Nurse Pru le Cuirot left her home in Jersey to care for injured soldiers at Ashbury Manor, Dorset. She wanted to do her bit but she never expected to meet Canadian pilot, Jack Garland, so unlike any man she has ever met. When Jack is sent on a dangerous mission and vanishes, Pru is forced to make an agonising decision…one that will haunt them both forever.

At 97 years old, Winnifred Ellis knows she doesn’t have much time left, and it is almost a relief to realize that once she is gone, the truth about her shameful past will die with her. But when her great-grandson Jamie, the spitting image of her dear late husband, asks about his family tree, Winnifred can’t lie any longer, even if it means breaking a promise she made so long ago....

London, 1935. Ruby is always hungry, but she will go without if it means her young brothers can eat. 1930s Bermondsey might be called the larder of London, with its pie, pickle and jam factories, but for the poor working classes, starvation is often only a heartbeat away. When Ruby's neighbour suggests she ought to go to the Methodist Mission for free food, Ruby knows her father will be furious, but she has no other option.

Margot and her adoring husband, Jochen, are madly in love. But in Nazi Germany, being in love is the most dangerous act of all. A marriage between Margot, a German woman, and Jochen, a Jewish man, is forbidden under Hitler’s tyrannical rule. Yet when friends cross the street to avoid Margot and her own mother rejects her, she stays loyal to Jochen. When the Gestapo break into their home and steal their belongings, she remains steadfast. And when soldiers arrest Jochen, imprisoning him with countless others, Margot will stop at nothing to save him.