When Betty Palmer's sister dies under suspicious circumstances whilst landing her Tiger Moth, Betty and three other women pilots of the Air Transport Auxiliary in WWII England unite to discover who killed her and why. Estranged from her family, Penny Blake wants simply to belong. American Doris Winter, running from a personal tragedy, yearns for a new start. Naturally shy Mary Whitworth-Baines struggles to fit in. Together, though, they are a force to be reckoned with as they face the mystery that confronts them.

Following a humiliating experience involving the man she thought she’d marry, Iris Tredwick signs up to the Wrens in order to escape and - hopefully - find ‘the right sort’ of man to please her mother. After a bumpy start, Iris manages to befriend outspoken Mary and dreamer Sally as they are sent to their first posting - in Orkney. There she meets engineer Rob, whose flirtatious nature both charms and confounds straight-laced Iris. Much more appropriate for her is local doctor Stewart, if only she felt the same spark for him as she does for Rob.

September 1939. London is in blackout; war has been declared, but Dulcie Treadwell can think only of her heartbreak over American broadcaster, Glenn Reeves, who didn’t say goodbye before leaving for Berlin. Posted to Bletchley Park, Dulcie must concentrate on helping the war effort by cracking the German Enigma codes. The hours are long and the conditions tough, with little recognition from above. Until she breaks her first code. Dulcie finally settles into something she was born to do. But just as she’s proving her worth, a shocking betrayal leads to Dulcie’s brutal dismissal.

Lady Joanna Harcourt and her daughter, Sarah, discover their life of luxury at Goodwill House is over. Because with Lord Harcourt away fighting, the Harcourt girls are facing financial ruin. Lady Joanna is terrified of losing their home, but for Sarah, this means her dreams of becoming a doctor are over. Headstrong Sarah isn’t one to quit! War or not, she’s determined to find a way to save her home and follow her dreams - whatever it takes.

Liverpool, 1939. Bella Rogers thought her life was just starting; she can’t wait to get married to her childhood sweetheart, Bobby. But when war is declared and Bobby is called up, his letters stop, leaving Bella heartbroken and alone. Then her family is torn apart by tragedy when her angelic 5-year-old sister dies from a terrible illness. They can barely afford a funeral, but the church is packed with the entire community, devastated for their loss. Grieving for her sister and lost love, Bella finds a job at the Bryant and May factory, making matches.

When sixteen-year-old Sally is evacuated to the English south coast, she is terrified by what lies ahead of her. All she knows are the sights and sounds of London's East End – but Sally swallows her tears as they leave the familiar landmarks behind, knowing that she has to be a Grown-Up Girl and play mother to her six-year-old brother Ernie. Playing mother is nothing new for Sally – their real mother Florrie, a good-time girl, hasn't even come to the station to wave them off and Ernie, crippled at an early age by polio, is used to depending on his older sister.

At the heart of Dover's Market Square lies the Castle's Café. A bustling, homely place run by the formidable Nellie Castle and her six children. Since the scandalous birth of her beloved son 10 years before, Marianne, Nellie's eldest daughter, has preferred to stay in the kitchen, hidden away from the town gossips. Overcome with shame, Marianne has never revealed the name of Donny's father - not even to her own mother. But with World War II just around the corner, Marianne's past catches up with her. And suddenly the lives of the Castle family become a lot more complicated.

Bristol 1936. Jenny Crawford has resigned herself to a loveless marriage living hand to mouth with their two children. Like many others, husband Roy struggles to find work at the docks but a chance encounter turns the family's fortunes around. Not only does he get regular work but they are also allocated a council house on Coronation Close on the outskirts of the city. Jenny and the children are overjoyed, this is the fresh start she could only ever dream of. But trouble feels never too far away.

London, 1941. The Blitz. When a Royal Navy memo arrives at head office, requesting female recruits to sign up as motorcycle dispatch riders, delivering highly classified orders across the country, three women jump at the chance to sign up for the most dangerous jobs in London.

Billeted together in the attic of Latimer House—a place where secrets abound—Evelyn, Betty and Judith soon form a bond of friendship that carries them through the war. Because nothing is stronger than women united. Tucked away in the Buckinghamshire countryside, Latimer House, a grand country estate, stands proudly—a witness to some of greatest secrets of WW2.

At the height of World War II, the British Air Transport Auxiliary need help. A group of young women volunteer for action, but the perils of their new job don’t end on the tarmac. Things are tough in the air, but on the ground their abilities as pilots are constantly questioned. There is friction from the start between the new recruits. Spirited American Lizzie turns heads with her audacity, but few can deny her flying skills. She couldn’t be more different from shy, petite Ruby, who is far from diminutive in the sky. It falls to pragmatic pilot May to bring the women together.

Natty and Sean Wainwright have a rock-solid marriage and family. When their younger daughter falls ill on an overseas school trip, Natty rushes to her side. Luckily, Natty's best friend from college, Eve Dalladay, is visiting and offers to stay with Sean to lend a hand in the Wainwright household. But Natty returns home to find that Sean has fallen in love with Eve. Natty attempts to start anew, but Eve is there to knock her down again.

Lizzie Dixon's life feels as though it's fallen apart. Instead of the marriage proposal she was hoping for from her boyfriend, she is unceremoniously dumped, and her job is about to go the same way. So there's only one option: to go back home to the village she grew up in and to try to start again. Her best friend, Jemma, is delighted Lizzie has come back home, as she has just bought a little cafe, and Lizzie's sewing skills are just what she needs to help get it ready for the grand opening.

Brighton, 1938: Grace Kemp is pushed away by the family she has shamed. Rejected and afraid, she begins a new life as a nurse. But danger stalks the hospital too, and she'll need to be on her guard to avoid falling into familiar traps. And then there are the things she sees....Strange portents that have a way of becoming real.

Liverpool, 1941. Troubled by a dark family history and the pain of seeing her country torn apart by war, young June Lavender finds new hope when she takes a job at the Dr Barnardo's children's home. There she meets Lizzie McRae, a little girl who is the spitting image of June's sister, who died tragically years before. Against the matron's wishes, the two girls form an unexpected bond as June tries to get the traumatised orphan to speak.